A true breeding plant with red flowers and pointy leaves is crossed to a true breeding plant with white flowers and round leaves. The F2 progeny are
451 pink heart-shaped
221 pink round
215 white heart-shaped
205 red heart-shaped
198 pink pointy
113 white round
111 red pointy
105 white pointy
98 red round
1. Figure out the plausible mendelian segregation for this cross—justify this using mendelian concepts
2. Test to see what happens
Solution
If we have a two loci, two alleles trait we have nine possible genotypes. If each possible genotype has a phenotype then we have nine phenotypes. Since we see nine phenotypes we hypothesize that this is the case.
So we hypothesize two loci, two alleles each with both loci co-dominant. In this case the segregation is
1:2:1:2:4:2:1:2:1
(1:2:1 times 1:2:1)
reorganizing the phenotypes and assigning hypothesized genotypes
111 red pointy AABB
198 pink pointy AaBB
105 white pointy aaBB
205 red heart-shaped AABb
451 pink heart-shaped AaBb
215 white heart-shaped aaBb
98 red round AAbb
221 pink round Aabb
113 white round aabb
|
|
observed |
expected |
0-e |
(0-e)^2 |
(0-e)^2/e |
|
|
111 |
107 |
3.7 |
13.598 |
0.12671 |
|
|
198 |
215 |
-17 |
276.39 |
1.28778 |
|
|
105 |
107 |
-2 |
5.3477 |
0.04983 |
|
|
205 |
215 |
-10 |
92.641 |
0.43164 |
|
|
451 |
429 |
22 |
473.06 |
1.10207 |
|
|
215 |
215 |
0.4 |
0.1406 |
0.00066 |
|
|
98 |
107 |
-9 |
86.723 |
0.80813 |
|
|
221 |
215 |
6.4 |
40.641 |
0.18936 |
|
|
113 |
107 |
5.7 |
32.348 |
0.30143 |
|
total |
1717 |
|
|
|
|
|
|
|
|
|
X2 |
4.29761 |
|
|
|
|
|
|
|
|
|
|
|
|
p-value |
0.8291 |
Fail to reject and I conclude: There is not enough evidence to conclude that the ratio is different from
1:2:1:2:4:2:1:2:1
Let’s do another
A true breeding plant with red flowers and pointy leaves is crossed to a true breeding plant with white flowers and round leaves. The F2 progeny are
316 red pointy
98 red round
221 pink round
649 pink pointy
320 white pointy
113 white round
1. Propose a plausible mendelian segregation
2. Test it
Solution
If we have a two loci two alleles trait we have nine possible genotypes. If one locus is co-dominant and one locus is dominant then we have six phenotypes. Since we see six phenotypes we hypothesize that this is the case. So we propose two loci, two alleles, one locus has a dominant allele, the other locus has co-dominant alleles. In this case the segregation is
3:6:3:1:2:1
(3:1 times 1:2:1)
Reordering the phenotypes
316 red pointy AABB /AABb
649 pink pointy AaBB/ AaBb
320 white pointy aaBB /aaBb
98 red round AAbb
221 pink round Aabb
113 white round aabb
|
o |
e |
0-e |
(0-e)^2 |
(0-e)^2/e |
|
316 |
322 |
-6 |
35 |
0.11 |
|
649 |
644 |
5.1 |
26 |
0.041 |
|
320 |
322 |
-2 |
4 |
0.012 |
|
98 |
107 |
-9 |
87 |
0.808 |
|
221 |
215 |
6.4 |
41 |
0.189 |
|
113 |
107 |
5.7 |
32 |
0.301 |
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
X2 |
|
1.461 |
|
|
|
|
|
|
|
|
|
pvalue |
0.92 |
|
Fail to reject and I conclude: There is not enough evidence to conclude that the ratio is different from 3:6:3:1:2:1.
Example
A wild legume with white flowers and long pods is crossed to one with purple flowers and short pods. The F1 flowers are allowed to self fertilize and the F2 generation has 301 long purple, 99 short purple, 612 long pink, 195 short pink, 295 long white, 98 short white
What is the mendelian segregation and test to see whether this follows.
Building the hypothesis:
PPll x ppLL p1xp2
PpLl F1
F2 genotypes
PPLL 2 PpLL ppLL
2 PPLl 4 PpLl 2 ppLl
PPll 2 Ppll ppLL
I have nine genotypic categories but now 6 phenotypic categories so I need to figure out what the mapping is between the genotype and the phenotype…
Long seems dominant to short. White and Purple appear to be co-dominant with pink as the heterozygote.
I hypothesize this is the relationship between genotype and phenotype
Genotype Phenotype Exp
PPLL+2PPLl purple long 3/16
PPll purple short 1/16
2PpLL+4PpLl pink long 6/16
2Ppll pink short 2/16
ppLL+2ppLl white long 3/16
ppll white short 1/16
Can also get the ratio by multiplying
1:2:1 by 3:1!
The null hypothesis is : 3:1:6:2:3:1
|
o |
e |
o-e |
(o-e)2 |
(o-e)2/e |
|
301 |
300 |
1 |
1 |
0.003333 |
|
99 |
100 |
-1 |
1 |
0.01 |
|
612 |
600 |
12 |
144 |
0.24 |
|
195 |
200 |
-5 |
25 |
0.125 |
|
295 |
300 |
-5 |
25 |
0.083333 |
|
98 |
100 |
-2 |
4 |
0.04 |
|
|
|
|
|
|
|
1600 |
|
|
X2 |
0.501667 |
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
|
p |
0.992062 |
Fail to reject the null
There is not enough evidence to conclude that the ratio is different from 3:1:6:2:3:1.